Suborder
Axonophora Frech, 1897
Selection of related publications
Maletz, J., Ahlberg, P. 2021. Dapingian to lower Darriwilian (Middle Ordovician) graptolite biostratigraphy and correlation of the Krapperup drill core, Scania, Sweden.  GFF   143,   1,   16-39. DOI:10.1080/11035897.2020.1822439
Maletz, J., Ahlberg, P., Lundberg, F. 2020. Ordovician graptolite biostratigraphy of the Röstånga-2 drill core (Scania, southern Sweden).  GFF   142,   3,   206-222. DOI:10.1080/11035897.2020.1739743
Loydell, D. K., Walasek, N. 2020. Two new species of graptolite from the Telychian (upper Llandovery, Silurian) of Kallholn, Dalarna, Sweden.  GFF   142,   2,   154-157. DOI:10.1080/11035897.2019.1686419
Kozłowska, A., Bates, D., Zalasiewicz, J., Radzevičius, S. 2019. Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania .  Zootaxa    4568,   3,   435. DOI:10.11646/zootaxa.4568.3.2
Štorch, P., Manda, Š. 2019. Little known Homerian (lower Silurian) graptolites from Kosov quarry near Beroun, the Czech republic.  Fossil Imprint   75,   1,   44-58. DOI:10.2478/if-2019-0003
Maletz, J. 2019. Second Revision, Chapter 22: Suborder Axonophora, Infraorder Diplograptina I (Diplograptidae, Lasiograptidae): Introduction, morphology, and systematic descriptions.  Treatise Online 127, pp. 1-19.   The University of Kansas.  DOI:10.17161/to.v0i0.13265
Kozłowska, A. 2016. A new generic name, Semigothograptus, for Gothograptus? meganassa Rickards & Palmer, 2002, from the Silurian post-lundgreni Biozone recovery phase, and comparative morphology of retiolitids from the lowermost upper Homerian (upper Wenlock).  Zootaxa    4208,   6,   534-546. DOI:10.11646/zootaxa.4208.6.2
Männik, P., Tinn, O., Loydell, D. K., Ainsaar, L. 2016. Age of the Kalana Lagerstätte, early Silurian, Estonia.  Estonian Journal of Earth Sciences   65,   2,   105-114. DOI:10.3176/earth.2016.10
Goldman, D., Nõlvak, J., Maletz, J. 2015. Middle to Late Ordovician graptolite and chitinozoan biostratigraphy of the Kandava-25 drill core in western Latvia.  GFF   137,   3,   197-211. DOI:10.1080/11035897.2015.1021375
Männik, P., Loydell, D. K., Nestor, V., Nõlvak, J. 2015. Integrated Upper Ordovician–lower Silurian biostratigraphy of the Grötlingbo-1 core section, Sweden.  GFF   137,   3,   226-244. DOI:10.1080/11035897.2015.1042032
Maletz, J., Steiner, M. 2015. Graptolites (Hemichordata, Pterobranchia) preservation and identification in the Cambrian Series 3.  Palaeontology   58,   6,   1073–1107. DOI:10.1111/pala.12200
Maletz, J., Ahlberg, P., Suyarkova, A., Loydell, D. K. 2014. Silurian graptolite biostratigraphy of the Röstånga-1 drill core, Scania – a standard for southern Scandinavia.  GFF   136,   1,   175-178. DOI:10.1080/11035897.2013.865665
Bergström, S. M., Eriksson, M. E., Young, S. A., Ahlberg, P., Schmitz, B. 2014. Hirnantian (latest Ordovician) d13C chemostratigraphy in southern Sweden and globally: a refined integration with the graptolite and conodont zone successions.  GFF   136,   2,   355-386. DOI:10.1080/11035897.2013.851734
Kozłowska, A., Radzevičius,  S. 2013. Upper Homerian and Gorstian (Silurian) Retiolitidae (Graptolithina) from Lithuania and Latvia.  Memoirs of the Association of Australasian Palaeontologists   44,      11-23.
Radzevičius, S. 2013. Silurian graptolite biozones of Lithuania: present and perspectives.  Geologija   55,   2(82),   41-49. DOI:10.6001/geologija.v55i2.2757
Kozłowska, A., Bates, D. E. B. 2013. Kirkigraptus, a new retiolitid graptolite from Poland.  Acta Palaeontologica Polonica   53,   1,   105–112. DOI:10.4202/app.2008.0107
Loydell, D. K. 2012. Graptolite biozone correlation charts.  Geological Magazine   149,   1,   124–132. DOI:10.1017/S0016756811000513
Paškevičius, J., Klimantavičius, V., Radzevičius, S. 2012. Lithostratigraphy, graptolites and brachiopods communities of the Ludlow (Silurian) of the Eastern slope of the Baltic Syneclise.  Geologija   54,   3,   75-88. DOI:10.6001/geologija.v54i3.2515
Urbanek, A., Radzewičius, S., Kozłowska, A., Teller, L. 2012. Phyletic evolution and iterative speciation in the persistent Pristiograptus dubius lineage.  Acta Palaeontologica Polonica   57,   3,   589-611. DOI:10.4202/app.2010.0070
Maletz, J., Ahlberg, 2011. Darriwilian (Ordovician) graptolite faunas and provincialism in the Tøyen shale of the Krapperup drill core (Scania, Southern Sweden).  Ordovician of the World, pp. 327-332.   © Instituto Geológico y Minero de España. 
Maletz, J. 2011. The identity of the Ordovician (Darriwillian) graptolite Fucoides dentatus Brogniart, 1828.  Palaeontology   54,   4,   851-865. DOI:10.1111/j.1475-4983.2011.01068.x
Maletz, J. 2011. The proximal development of the Middle Ordovician graptolite Skanegraptus janus from the Krappegrup drill core of Scania, Sweden.  GFF   133,   1-2,   49-56. DOI:10.1080/11035897.2011.564293
Maletz, J., Egenhoff, S., Böhme, M., Asch, R., Borowski, K., Höntzsch, S., Kirsch, M., Werner, M. 2011. A tale of both sides of Iapetus – upper Darriwilian (Ordovician) graptolite faunal dynamics on the edges of two continents.  Canadian Journal of Earth Sciences   48,   5,   841-859. DOI:10.1139/e10-105
Pärnaste, H., Popp, A. 2011. First record of Telephina (Trilobita) from the Ordovician of northeastern Estonia and its stratigraphical implications.  Estonian Journal of Earth Sciences   60,   2,   83-90. DOI:10.3176/earth.2011.2.02
Loydell, D. K., Nestor, V., Männik, P. 2010. Integrated biostratigraphy of the lower Silurian of the Kolka-54 core, Latvia.  Geological Magazine   147,   2,   253-280. DOI:10.1017/S0016756809990574
Kozłowska, A., Lenz, A., Melchin, M. 2009. Evolution of the Retiolitid Neogothograptus (Graptolithina) and Its New Species from the Upper Wenlock of Poland, Baltica.  Acta Palaeontologica Polonica   54,   3,   423-434. DOI:10.4202/app.2008.0022
Suyarkova, A. A., Koren', T. N. 2009. Advances in the Ordovician graptolite biostratigraphy of the St Petersburg area, Russia.  Estonian Journal of Earth Sciences   58,   1,   3-9. DOI:10.313176/earth.2009.1.01
Mitchell, C. E., Maletz, J., Goldman, D. 2009. What is Diplograptus?.  Bulletin of Geosciences   84,   1,   27-34. DOI:10.3140/bull.geosci.1109
Loydell, D. K., Maletz, J. 2009. Isolated graptolites from the Lituigraptus convolutus biozone (Silurian, Llandovery) of Dalarna, Sweden.  Palaeontology   52,   2,   273-296. DOI:10.1111/j.1475-4983.2008.00843.x
Radzevičius, S., Kiipli, T., Kallaste, T., Motuza, V., Jeppsson, L. 2008. Wenlock bentonites in Lithuania in the frame of graptolite stratigraphy.  The Seventh Baltic Stratigraphical Conference, 15-22 May 2008, Estonia. Abstracts, pp. 56-56. 
Nõlvak, J., Goldman, D. 2007. Biostratigraphy and taxonomy of three-dimensionally preserved nemagraptids from the Middle and Upper Ordovician of Baltoscandia.  Journal of Paleontology   2007,   2,   254-260. DOI:10.1666/0022-3360(2007)81[254:BATOTP]2.0.CO;2
Loydell, D. K., Jeppsson, L. 2006. Graptolites from the Lower and Upper Visby formations of NW Gotland.  GFF   128,   2,   159-160. DOI:10.1080/11035890601282159
Bates, D. E. B., Kozłowska, A., Maletz, J., Kirk, N. H., Lenz, A. 2006. The Silurian retiolitid graptolite Plectograptus: New observations and new species.  Acta Palaeontologica Polonica   51,   3,   525-540.
Radzevičius, S. 2006. Late Wenlock biostratigraphy and the Pristiograptus virbalensis group (Graptolithina) in Lithuania and the Holy Cross Mountains.  Geological Quarterly   50,   3,   333-344.
Loydell, D. K., Nestor, V. 2006. Isolated graptolites from the Telychian (Upper Llandovery, Silurian) of Latvia and Estonia.  Palaeontology   49,   3,   585-619. DOI:10.1111/j.1475-4983.2006.00550.x
Loydell, D. K., Nestor, V. 2005. Integrated graptolite and chitinozoan biostratigraphy of the upper Telychian (Llandovery, Silurian) of the Ventspils D-3 core, Latvia.  Geological Magazine   142,   4,   369-376. DOI:10.1017/S0016756805000531
Nõlvak, J., Goldman, D. 2004. Distribution of Nemagraptus in the East Baltic Ordovician.  WOGOGOB-2004. Conference Materials. Abstracts and Field Guide Book, pp. 75-76.   Tartu University Press. 
Loydell, D. K., Maletz, J. 2004. The Silurian graptolite genera Streptograptus and Pseudostreptograptus.  Journal of Systematic Palaeontology   2,   2,   65-93. DOI:10.1017/S1477201904001117
Loydell, D. K., Männik, P., Nestor, V. 2003. Integrated biostratigraphy of the lower Silurian of the Aizpute-41 core, Latvia.  Geological Magazine   140,   2,   205-229. DOI:10.1017/S0016756802007264
Põldvere, Anne (ed) 2003. Ruhnu (500) drill core.  Estonian Geological Sections Bulletin   5,      1-76.  Geological Survey of Estonia. 
Pålsson, C. 2002. Upper Ordovician graptolites and biostratigraphy of the Röstånga 1 core, Scania, S. Sweden.  Bulletin of the Geological Society of Denmark   49,   1,   9-23. DOI:10.37570/bgsd-2003-49-02
Kozłowska-Dawidziuk, A. 2002. Agastograptus, a synonym of Plectograptus (Retiolitidae, Graptolithina).  Acta Palaeontologica Polonica   47,   3,   459-467.
Pålsson, C. 2001. Graptolites from the Upper Ordovician Dicranograptus clingani Zone at Järrestad, Scania, southern Sweden.  GFF   123,   4,   217-224. DOI:10.1080/11035890101234217
Kozłowska-Dawidziuk, A. 2001. Phylogenetic relationships within the Retiolitidae (Graptolithina) and a new genus, Cometograptus.  Lethaia   34,   1,   84 - 96. DOI:10.1080/002411601300068314
Radzevičius, S., Paškevičius, J. 2000. Pristiograptids (Graptoloidea) and their adaptive types of the Wenlock (Silurian) in Lithuania.  Geologija   32,      88-109.
Loydell, D. K., Kaljo, D., Männik, P. 1998. Integrated biostratigraphy of the lower Silurian of the Ohesaare core, Saaremaa, Estonia.  Geological Magazine   135,   6,   769-783. DOI:10.1017/S0016756898001423
Urbanek, A. 1997. Late Ludfordian and early Přidoli monograptids from the Polish Lowland.  Palaeontologia Polonica   56,      87-231.
Kozłowska-Dawidziuk, A. 1997. Retiolitid graptolite Spinograptus from Poland and its membrane structures.  Acta Palaeontologica Polonica   42,   3,   505-518.
Raukas, A., Teedumäe, A. (eds) 1997. Geology and Mineral Resources of Estonia. pp. 1-436.  Estonian Academy Publishers. 
Maletz, J. 1997. Graptolites from the Nicholsonograptus fasciculatus and Pterograptus elegans Zones (Abereiddian, Ordovician) of the Oslo Region, Norway.  Greifswalder Geowissenschaftliche Beiträge   4,      5-98.
Koren', T., Bjerreskov, M. 1997. Early Llandovery monograptids from Bornholm and the southern Urals: taxonomy and evolution.  Bulletin of the Geological Society of Denmark   44,   1,   1-43. DOI:10.37570/bgsd-1998-44-01
Kozłowska-Dawidziuk, A. 1995. Silurian retiolitids of the East European Platform.  Acta Palaeontologica Polonica   40,   3,   261-326.
Mitchell, C. E., Maletz, J. 1995. Proposal for adoption of the base of the Undulograptus austrodentatus Biozone as a global Ordovician stage and series boundary level.  Lethaia   28,   4,   317-331. DOI:10.1111/j.1502-3931.1995.tb01822.x
Maletz, J. 1995. The Middle Ordovician (Llanvirn) graptolite succession of the Albjära core (Scania, Sweden) and its implication for a revised biozonation.  Zeitschrift für Geologische Wissenschaften   23,   3,   249-259.
Jaanusson, V. 1995. Confacies differentiation and upper Middle Ordovician correlation in the Baltoscandian basin.  Proceedings of the Estonian Academy of Sciences. Geology   44,   2,   73-86. DOI:10.3176/geol.1995.2.01
Kozłowska-Dawidziuk, A. 1991. Agastograptus from the Mulde beds of Gotland.  Acta Palaeontologica Polonica   36,   2,   143-149.
Kozłowska-Dawidziuk, A. 1990. The genus Gothograptus (Graptolithina) from the Wenlock of Poland.  Acta Palaeontologica Polonica   35,   3-4,   191-209.
Bassett, M. G., Kaljo, D., Teller, L. 1989. The Baltic region.  National Museum of Wales, Geological Series   9,      158-170.  National Museum of Wales. 
Obut, A. M., Sennikov, N. V. 1987. Oepikograptus, a pecuilar genus of Middle Ordovician diplograptids (graptolites).  Annual of the All-union Paleontological Society. Volume XXX, pp. 192-198.   Nauka. 
Ulst, R. 1986. Osobennosti stroeniâ proksimpli u nekotoryh llandoverijskih graptolitov Latvii.  Biofacies and fauna of the Silurian and Devonian basins of the Baltic, pp. 22-24.   Zinatne. 
Williams, S. H., Bruton, D. L. 1984. The Caradoc- Ashgill boundary in the central Oslo region and associated graptolite faunas ( Norway).  Norsk Geologisk Tidsskrift   63,   2,   147-191.
Ulst, R. Ž., Gailite, L. K., Springis, T. K. 1984. Lithostratigraphic subdivision of the subsurface Ordovician rocks of the Jelgeva Depression.  Stratigrafiâ drevnepaleozojskih otloženij Pribaltiki, pp. 63-76.   Eesti NSV Teaduste Akadeemia. 
Kaljo, D., Paškevičius, J., Ulst, R. 1984. Graptolite zones of the East Baltic Silurian.  Stratigrafiâ drevnepaleozojskih otloženij Pribaltiki, pp. 94-118.   Eesti NSV Teaduste Akadeemia. 
Obut, A. M., Zaslavskaya, N. M. 1983. The families of retiolitids and their phylogenetic relationships.  Transactions of the Institute of Geology and Geophysics   538,      103-113.  Nauka. 
Howe, M. P. A. 1982. The Lower Silurian graptolites of the Oslo region.  Field Meeting, Oslo Region, 1982, pp. 21-32. 
Paškevičius, J. 1979. Biostratigraphy and Graptolites of the Lithuanian Silurian. pp. 1-267.  Mokslas. 
Urbanek, A. 1979. Sabellidites.  Upper Precambrian and Cambrian Paleontology of East-European Platform, pp. 87-92.   Nauka. 
Männil, Ralf 1976. Distribution of graptoloids in the Ordovician carbonate rocks of the East Baltic area.  Graptolites and Stratigraphy, pp. 105-118.   Academy of Sciences of Estonian SSR. Institut of Geology. 
Paškevičius, J. 1976. On some new Llandoverian diplograptids of the Eastern Baltic.  Graptolites and Stratigraphy, pp. 140-151.   Academy of Sciences of Estonian SSR. Institut of Geology. 
Tsegelnjuk, P. D. 1976. Late Silurian and Early Devonian monograptids from the south-western margin of the East-European platform.  Upper Precambrian and Lower Paleozoic paleontology and stratigraphy of the south-western part of the East European Platform, pp. 91-133.   Naukova Dumka. 
Ulst, R. 1975. Graptolites from Upper Ordovician Fjäcka Member of Middle Baltic.  The Fauna and Stratigraphy of Paleozoic and Mesozoic of Baltic and Byelorussia, pp. 69-84.   Mintis. 
Ulst, R. Ž. 1974. The early sequence of Pristiograpts in conterminous deposits of Wenlock and Ludlow of the Middle Pribaltic.  Graptolites of the USSR, pp. 105-122.   Nauka. 
Paškevičius, J. 1974. Graptolites and zonal dismemberment of Ludlow deposits in Pribaltic.  Graptolites of the USSR, pp. 122-134.   Nauka. 
Paškevičius, J. 1973. Biostratigraphy, correlation and graptolites of the Ordovician and Silurian deposites of the South East Baltic.  kand töö autoreferaat, pp. 1-63. 
Rõõmusoks, A. 1970. Stratigraphy of the Viruan Series (Middle Ordovician) in Northern Estonia, I. pp. 1-346.  Valgus. 
Urbanek, A. 1970. Neocucullograptinae n. subfam. (Graptolithina)- their evolutionary and stratigraphic bearing.  Acta Palaeontologica Polonica   15,   2-3,   164-388.
Kaljo, D., Vingisaar, P. 1969. On the sequence of the Raikküla stage in southernmost Estonia.  Eesti NSV Teaduste Akadeemia Toimetised. Keemia, Geoloogia / Известия Академии наук Эстонской ССР. Химия. Геология   18,   3,   270-277. DOI:10.3176/chem.geol.1969.3.10
Krandijevsky, V. S. 1968. Граптолиты силура Волыно-Подолии.  Paleontology and Stratigraphy of the Lower Paleozoic of Volyn-Podola, pp. 26-62.   Naukova Dumka. 
Gailite, L. K., Rybnikova, M. V., Ulst, R. Ž. 1967. Stratigrafiâ, fauna i usloviâ obrazovaniâ silurijskih porod Srednej Pribaltiki. pp. 1-304.  Zinatne. 
Urbanek, A. 1966. On the morphology and evolution of the Cucullograptinae (Monograptidae, Graptolithina).  Acta Palaeontologica Polonica   11,   3-4,   292-544.
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Jaanusson, V., Skoglund, R. 1963. Graptoloids from the Viruan (Ordovician) Dalby and Skagen Limestones of Västergötland.  GFF   85,   3,   341-357. DOI:10.1080/11035896309452966
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List of species
1. Akidograptus ascensus Davies, 1929 | Rhuddanian
5. Amplexograptus pulchellus (Hadding, 1915) | Upper Ordovician
6. Amplexograptus tubulus Berry, 1964 | Ordovician
7. Appendispinograptus supernus (Elles et Wood, 1906) | Upper Ordovician
8. Archiclimacograptus angulatus (Bulman, 1953) | Middle Ordovician
10. Archiclimacograptus caelatus (Lapworth, 1875) | Darriwilian
11. Archiclimacograptus decoratus (Harris et Thomas, 1935) | Darriwilian
12. Archiclimacograptus marathonensis (Clarkson, 1963) | Middle Ordovician
13. Archiclimacograptus micidus (Berry, 1964) | Middle Ordovician
14. Archiclimacograptus modestus (Ruedemann, 1908) | Darriwilian
15. Archiclimacograptus osloensis Goldman, Nõlvak et Maletz, 2015 | Darriwilian
16. Archiclimacograptus riddellensis (Harris, 1924) | Darriwilian
17. Archiclimacograptus sebyensis (Jaanusson, 1960) | Ordovician
18. Archiclimacograptus skagensis (Jaanusson et Skoglund, 1963) | Sandbian
20. Atavograptus atavus (Jones, 1909) | Llandovery
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Classification
| Organism group | Biota | 
| Kingdom | Animalia | 
| Phylum | Hemichordata | 
| Class | Pterobranchia | 
| Subclass | Graptolithina | 
| Order | Graptoloidea | 
| Suborder | Axonophora | 
| Infraorder | Diplograptina | 
| Neograptina | |
| Superfamily | Retiolitoidea | 
| Family | Climacograptidae | 
| Dicranograptidae | |
| Lasiograptidae | |
| Monograptidae | |
| Subfamily | Orthograptinae | 
| Suborder | Dichograptina | 
| Glossograptina | |
| Graptodendroidina | |
| Family | Cryptograptidae | 
| Cyrtograptidae | |
| Dicaulograptidae | |
| Glyptograptidae | |
| Linograptidae | |
| Genus | Corymborgaptus |