Genus

Protovirgularia McCoy, 1850

Taxon description

Uchman, 1998

Emended  diagnosis : Horizontal or subhorizontal cylindri­cal trace fossil, trapezoidal, almond or triangular in crosssection, distinctly or in distinctly bilobate . Internal structure can be preserved: it is formed by successive pads of sedi­ment which can be expresed on the exterior as ribs. The ribs are arranged in a chevron -like biserial pattern along the external or internal dorsal part. Exterior smooth mantle coverin g the structure and /or oval m ound -like terminations of the trace fossil can be present.

Luo & Shi, 2017

Diagnosis. Horizontal to subhorizontal keel-like bilobated trails, composed of chevronate ribs arranged to form a carinate pattern. Trail cross-section trapezoidal, amygdaloidal to triangular, preserved as both convex/concave epirelief and positive hyporelief (emended from Han & Pickerill 1994 and Uchman 1998).

Remarks. The morphological features of Protovirgularia are similar to several other horizontal trails, such as Cruziana, Scolicia,Nereites, and Psammichnites. The bilobate Cruziana has numerous scratch markings on both sides of the furrow, a feature never observed in Protovirgularia. Bilaterally symmetrical Scolicia lacks chevronate ribs that are typically developed in Protovirgularia. Nereites has a wider median furrow rather than a central string, and the closely spaced lobes on either side of the furrow never chevronate when compared with Protovirgularia. The lack of a median dorsal structure (e.g., straight ridge/groove) distinguishes Protovirgularia from Psammichnites.

Selection of related publications
Uchman, A., Hanken, N.-M., Binns, R. 2005. Ordovician Bathyal Trace Fossils From Metasiliciclastics in Central Norway and Their Sedimentological and Paleogeographical Implications. Ichnos 12, 2, 105-133. DOI:10.1080/10420940590914534
Hecker, R. F. 1957. Vedenie v paleoèkologiû. pp. 1-124. Gosudarstvennoe naučno-tehnicheskoe izdatel'stvo literatury po geologii i ohrane nedr.
Eichwald, E. 1860. Lethaea Rossica ou Paléontologie de la Russie. Premier volume. Ancienne Période en deux sections. pp. XIX+1–1657. E. Schweizerbart. DOI:10.5962/bhl.title.52391
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Classification
Organism groupBiota
Ichnofossil groupIchnofossils
Bioturbation trace fossils
Trackways and scratch imprints
OrderBiformitidae
FamilyArenicolitidae
Arthrophycidae
Asteriacitidae
Rhizocoralliidae
Rosselichnidae
Siphonichnidae
GenusAcanthorhaphe
Aglaspidichnus
Agrichnium
Altichnus
Amanitichnus
Amphorichnus
Ancorichnus
Angulichnus
Arachnomorphichnus
Arachnostega
Arborichnus
Archaeonassa
Arcuatichnus
Arenituba
Artharia
Asaphoidichnus
Asterichnus
Asterosoma
Astropolichnus
Aulichnites
Avipeda
Avolatichnium
Balanoglossites
Baroccoichnites
Bathichnus
Beaconites
Bergaueria
Bestiopeda
Bicavichnites
Bichordites
Bifasciculus
Bifungites
Bilinichnus
Bolonia
Bornichnus
Bromleyia
Bunyerichnus
Caletichnus
Calycraterion
Camborygma
Capodistria
Caprionichnius
Cardioichnus
Caridolites
Catenarichnus
Catenichnus
Chagrinichnites
Cheiichnus
Chomatichnus
Chondrites
Circulichnis
Cladichnus
Clematischnia
Climactichnites
Cochlichnus
Compaginatichnus
Conichnus
Conostichus
Cruziana
Ctenopholeus
Curvolithus
Cycloichnus
Cylindricum
Dactyloidites
Deliatynichnis
Dendrorhaphe
Dendrotichnium
Dictyodora
Didymaulichnus
Didymaulyponomos
Diplopodichnus
Echinospira
Egoreichnis
Elingua
Enigmatichnus
Entradichnus
Epibaion
Estrellichnus
Falcichnites
Fimbritubichnus
Foveichnus
Fucoides circinatus
Fuersichnus
Funalichnus
Furculosus
Fustiglyphus
Glossifungites
Glyphichnus
Gordia
Guanshanichnus
Gyrochorte
Gyrolithes
Gyrophyllites
Haentzschelinia
Halimedides
Halopoa
Hartsellea
Heimdallia
Helicodromites
Heliochone
Helminthoida
Helminthoidichnites
Helminthopsis
Herpystezoum
Hondichnus
Hormosiroidea
Ilmenichnus
Imponoglyphus
Isopodichnus
Jamesonichnites
Katbergia
Keilorites
Kimberichnus
Koptichnus
Korymbichnus
Kouphichnium
Labyrintichnus
Lajasichnus
Lamellaeichnus
Lamonte
Lanicoidichna
Lennea
Lepeichnus
Lepidenteron
Liholites
Lingulichnites
Lingulichnus
Lithoplaision
Lockeia
Loloichnus
Lophoctenium
Lorenzinia
Macaronichnus
Maiakarichnus
Mammillichnis
Margaritichnus
Megagrapton
Mermia
Microspherichnus
Minterichnus
Monocraterion
Monofungites
Multilamella
Multina
Multipodichnus
Musculopodus
Naviculichnium
Nenoxites
Neoeione
Neonereites
Nereites (trace fossil)
Nummipera
Oikobesalon
Oldhamia
Olenichnus
Oniscoidichnus
Ophiomorpha
Oravaichnium
Palaeophycus
Paleodictyon
Paracanthorhaphe
Parahaentzschelina
Parapsammichnites
Parataenidium
Paratisoa
Parmaichnus
Parundichnia
Pentichnus
Petalichnus
Phoebichnus
Pholeus
Phycosiphon
Pilichnus
Piscichnus
Plagiogmus
Plangtichnus
Planispiralichnus
Planolites
Polarichnus
Polykampton
Polykladichnus
Protopaleodictyon
Protospiralichnus
Protovirgularia
Speciesdichotoma
longespicata
mongraensis
obliterata
pennata
rugosa
triangularis
unrankednereitatum
GenusPsammichnites
Psilonichnus
Pteraspisichnis
Ptychoplasma
Radiichnus
Radionereites
Rhadhostium
Rotamedusa
Rusophycus
Saerichnites
Sagittichnus
Saportia
Scalaridomus
Schaubcylindrichnus
Scolecocoprus
Scolicia
Scotolithus
Scoyenia
Selenichnites
Shanwangichnus
Sinusichnus
Skolichnus
Skolithos
Sokolovichnites
Solemyatuba
Sphaerichnus
Spirocosmorhaphe
Spirodesmos
Spirophycus
Spongelimorpha
Squamichnus
Squamodictyon
Steinsfjordichnus
Stelloglyphus
Streptichnus
Sustergichnus
Svalbardichnus
Syringomorpha
Taenidium
Tambia
Taotieichnus
Taphrhelminthopsis
Taxichnites
Teichichnus
Ternavites
Tetraichnites
Thalassinoides
Torrowangea
Transexcrementum
Treptichnus
Triacliichnis
Trichichnus
Trichophycus
Trisulcus
Tuapseichnium
Tuberculichnus
Tubichnus
Tubotomaculum
Tylichnus
Undichna
Vagorichnus
Vermichnus
Virgaichnus
Volkichnium
Walcottia
Zoophycos
unrankedTrusheimichnus