Karethraichnus Zonneveld, Bartels, Gunnell et McHugh, 2016

Emended diagnosis.—Circular to subcircular non-penetrative and penetrative holes bored into osteic substrates. The boring that terminates within the substrate, forms a shallow, bowl-shaped pit. If the holes are deeper, they display a deeper shaft with a rounded to flattened terminus. Penetrative holes usually have straight or convex vertical edges.

Diagnosis: Circular to subcircular holes bored into a bone substrate. Holes may penetrate fully though the substrate or terminate within the bone. Penetrative holes may have straight or convex vertical margins. Non-penetrative pits terminate within the substrate as a shallow, bowl-shaped pit or as a deeper shaft with a rounded, blunt, or pointed terminus.

Macroboring; substrate osteic; tracemaker invertebrate