Ichnofossils
Species

Chondrites intricatus (Brongniart, 1823)

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Taxon description

Baucon et al., 2020c

Chondrites intricatus (Brongniart, 1828) has downward radiating straight branches; branching angle less than 45° (Uchman, 1998). Fu (1991) and Uchman (1999) placed Chondrites furcatus Sternberg, 1833 in synonymy with Chondrites intricatus, but Blissett and Pickerill (2004) have retained C. furcatus as a distinct ichnospecies;

Razzolini et al., 2016

Description. Three morphotypes A (the most common), B and C, are recognized. Morphotype A (Figs. 2C, 3B, 6A-B) is a system of tree-like branching, downward penetrating, markedly flattened tunnels. The width of the tunnels is constant in each specimen, but ranges from 0.2 to 0.6 mm overall. Branches form sharp angles. In most specimens, only second order branches occur; third-order branches are much rarer. Width of the burrow system ranges from 20 to 35 mm. The tunnels are filled with clayey material that is darker than the host rock and derives from the background sediments. Morphotype A penetrates up to 10 cm below the top of the marlstones. Morphotype B (Figs.4F,6C) occurs exclusively in fillings of Planolites beverleyensis as short, branched and unbranched, straight or slightly curved dark bars about 0.5 mm wide and up to 3 mm long. Morphotype C (Fig. 2D) is represented by branched tunnels of general geometry as in morphotype A, but tunnels are subdivided into segments that are less than 1 mm long (specimen no. 5081).

Remarks. Chondrites is a deep-tier trace fossil produced probably by surface ingestors (Kotake, 1991) that probably live in dysaerobic conditions as chemosymbiotic organisms (Seilacher, 1990; Fu, 1991).

Synonymy list
1823     Fucoides intricatus — Brongniart , pp. 311 , fig. 19:8
1828     Fucoides intricatus — Brongniart , pp. 59 , fig. 5:6-8
1896     Phycopsis intricata Brong, — Rothpletz , pp. 888 , fig. 22:7, 23:1
1902     Phycopsis arbuscula Fisher-Ooster — Rhotpletz , fig. 10b
1902     Phycopsis intricata Brongniart — Rothpletz , fig. 10c
1927     Chondrites intricatus Sternberg — Richter , fig. 8
1963     Chondrites intricatus — Müller , fig. 825
1977     Chondrites intricatus (Brogniart, 1823) — Książkiwicz , pp. 80 , fig. 4:5
1978     Chondrites intricatus — Kern , pp. 8246 , fig. 8B
1991     Chondrites intricatus (Brogniart, 1823) — Fu , pp. 18 , fig. 1E; 2A; Abb.10
1998     Chondńtes intricatus (Brongniart, 1828) — Gluszek , pp. 524 , fig. 4A, C
1998     Chondrites intricatus (Brogniart, 1823) — Głuszek , pp. 524 , fig. 4A,C
1999     Chondrites intricatus (Brogniart, 1823) — Uchman , pp. 88 , fig. 4:6-7; 5:1; 7:1-3, 5; text fig. 2
2001     Chondrites intricatus (Brogniart, 1823) — Uchman , pp. 10 , fig. 4:3; 6:4; 8:6
2003     Chondrites intricatus (Brogniart, 1823) — Leszczyński , pp. 111 , fig. 15:1, 16:2; 17:1; 25:2
2004     Chondrites intricatus — Leszczynski , pp. 41 , fig. 10, 12
2005     Chondrites intricatus (Brogniart, 1823) — Uchman et al. , pp. 60 , fig. 3B
2007     Chondrites intricatus (Brogniart, 1823) — Uchman , pp. 984-985 , fig. 2C-D; 3B, 4F, 6A-C
2011     Chondrites intricatus (Brogniart, 1823) — Gaillard , fig. 10:1
2012     Chondrites intricatus (Brogniart, 1823) — Uchman et al. , fig. 3
2012     Chondrites intricatus (Brogniart, 1823) — Uchman & Wetzel , fig. 2A
2016     Chondrites intricatus (Brogniart, 1823) — Ding et al. , pp. 379 , fig. 3A, B
2017     Chondrites intricatus — Knaust , fig. 5.34 a,b,d,e
2019     Chondrites intricatus — Toom et al. , fig. 4C
2020     Chondrites intricatus (Brogniart, 1823) — Baucon et al. , pp. 5 , fig. 2B
2022     Chondrites intricatus (Brogniart, 1823) — Šamánek et al. , pp. 774 , fig. 7A
2022     Chondrites intricatus (Brogniart, 1823) — Baucon et al. , pp. 6 , fig. 4B, 5A, B
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Classification
Organism groupBiota
Ichnofossil groupIchnofossils
Soft-sediment trace fossils
GenusChondrites
Speciesbollensis
affinis
caespitosus
intricatus
patulus
recurvus
stellaris
targionii