Kouphichnium Nopcsa, 1923

Diagnosis: Heteropodous trackway of great variability; complete trackway consisting of two kinds of track series: (1) two chevron-like series each of four oval, round, or bifid V-shaped impressions or scratches, forwardly directed; and (2) one pair of digitate or flabellar, toe-shaped or otherwise variable imprints; trackway with or without median drag ornament. All elements negative in epirelief and positive in hyporelief (modified after ROMANO & WHYTE 2003).